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    HYDROXYPROPYL DISTARCH GLYCEROL

    Explanation

         This modified starch was previously evaluated for an ADI for man
    by the Joint FAO/WHO Expert Committee on Food Additives in 1969, 1971
    and 1973 (see Annex I, Refs. 19, 26 and 29). Toxicological monographs
    were published in 1969, 1972 and 1974 (see Annex I, Refs. 20, 27 and
    30).

         Since the previous evaluation, additional data have become
    available and are summarized and discussed in the following monograph.
    The previously published monograph has been expanded and is reproduced
    in its entirety below.

    Introduction

         Modification is performed by the use of 0.1% epichlorohydrin and
    up to 10% propylene oxide. Cross-linking would be no greater than
    expected on modification with 0.1% epichlorohydrin. Substitution by
    treatment with 10% propylene oxide would probably add no more than
    15-20 ether linkages per 100 glucopyranose units.

    BIOLOGICAL DATA

    BIOCHEMICAL ASPECTS

         The in vitro digestibility of a hydroxypropyl distarch glycerol
    (degree of substitution 0.04) by pancreatin and porcine intestinal
    mucase was found to be 86% of that of the native starch (Leegwater,
    1971).

    TOXICOLOGICAL STUDIES

    Special studies on reproduction

         Groups each of 10 male and 20 female rats were fed the modified
    starch at a level of 10% in their diet for a period of 3 generations.
    Rats were mated (P1, F1 and F2) at weeks 12 and 20 after weaning.
    Each generation was mated twice and the litter from the second
    generation used to produce the next generation. No adverse effects
    were noted on fertility, number of pups per litter, body weight of
    young, mortality experience or resorption quotient that could be
    ascribed to treatment. The average relative caecum weight (filled) of
    females of the P and F2 generation was greater than controls, but
    empty caecum weight did not differ from controls. The relative caecum
    weight (filled and empty) of males of the F1 generation did not
    differ from controls. Growth of the F3b generation maintained on
    the test diet for 3 weeks was normal. Relative organ weights were
    comparable to control, with the exception of decreased thyroid weight

    and distinctly increased caecal weight (both filled and empty) in male
    pups, and increased empty caecal weight in female pups. Gross
    examination of organs at autopsy did not show any compound-related
    changes. Gross and microscopic examination of F3b rats did not show
    any compound-related changes (de Groot et al., 1974; Til et al.,
    1971b).

         In a modified 13-week subacute 1-generation reproduction study,
    hydroxypropyl distarch glycerol or cornstarch was fed at 10% of the
    diet to Charles River albino rats from 2 weeks prior to mating for the
    F1a generation until the sacrifice of the parental females for the
    caesarean delivery of the F1b generation (Olson & Ferrell, 1970).
    Twenty-five weanling F1a males and 25 weanling F1a females were
    randomly selected to be placed on a 13-week subacute dietary study.
    The reproduction indices for the F1a litter were comparable between
    the modified starch fed and unmodified starch groups. Skeletal
    evaluation of the F1b litter did not show any compound-related
    alternations. Organ weight, necropsy, and microscopic examination of
    the F1a pups did not show any compound-related effects. The modified
    starch-fed P females had elevated spleen and kidney weight ratios. The
    F1a generation fed for 13 weeks did not show any consistent effect
    which could be compound related. However, blood chemistry analyses
    showed elevated glucose for the females at the initial and 13-week
    intervals and for the males at 13 weeks. BUN was also slightly higher
    for both males and females at 13 weeks.

         Statistical analysis showed lower values for male terminal body
    weight, absolute thyroid and relative thyroid in the test animals as
    compared to controls, and higher values for female kidney weight and
    kidney ratio, and male kidney ratio than controls. No specific
    pathological alterations which could be attributed to the test
    procedure was seen in the animals maintained on either diet for 13
    weeks after weaning (Olson & Ferrell, 1970).

    Short-term studies

    Rat

         Groups of 10 male and 10 female rats were fed for 8 weeks on a
    diet containing 25 or 50% modified starch. Although the body weights
    of males at the higher level tested were slightly lower than those of
    the controls, the differences were not statistically significant. The
    faecal dry matter content was increased at both test levels in both
    sexes and there was considerable diarrhoea at the 50% level in both
    sexes. Slight diarrhoea occurred at the 25% level in both sexes.
    Caecal weights were increased in a dose-related manner at all levels
    in all test groups but no histological abnormalities could be detected
    in the caeca (de Groot & Spanjers, 1970).

         Groups of 25 male and 25 female rats were fed for 13 weeks on a
    diet containing 1 and 5% of modified starch or 5% of native starch.
    The general appearance and behaviour of test rats was comparable with
    the controls. One male and 1 female rat died at the lower test level.
    Growth, feed consumption, body weight, haematology at and 3 mouths,
    urinalysis at 1 and 3 months, and gross and histopathological findings
    were comparable in all groups (Knapp, 1967a).

         In another experiment, groups of 10 male and 10 female rats were
    fed for 90 days diets containing 0, 5, 10 or 30% modified starch. No
    differences between test and control animals were noted with regard
    to appearance, behaviour, growth, food consumption, haematology serum
    chemistry, or urinalysis. No definite diarrhoea occurred at any
    test level. The amounts of dry matter in the faeces did not vary
    significantly between controls and treated animals, only at the 30%
    level were the caecal weights increased in both sexes. Gross and
    histopathology were otherwise unremarkable in all groups.
    Histopathology of the enlarged caeca showed no recognizable
    abnormality (de Knecht-van Eekelen et al., 1970).

    Dog

         Three groups of 4 male and 4 female beagle dogs were given in
    their diet 1 and 5% of modified starch or 5% of native starch for 13
    weeks. Haematological studies, blood sugar, blood urea, serum GPT and
    alkaline phosphatase as well as urinalysis were comparable for all
    groups. Appearance and behaviour were normal. Body weight changes were
    unrelated to the administration of the test substance. Gross and
    histopathology showed no consistent abnormalities. Organ weights of
    thyroid, heart, liver, spleen, kidneys, ardenals and testes showed no
    test-related changes (Knapp, 1967b).

         In a second study with dogs (Olson & Voelker, 1970), groups of 4
    male and 4 female beagles were maintained on diets containing 10%
    hydroxypropyl distarch glycerol or cornstarch for a period of 13
    weeks. Food consumption and weight gain were comparable between the 2
    groups. Haematology, blood chemistry and urinalysis of the test and
    control groups were carried out at the beginning of the study and at
    week 13. There were no significant differences between the groups, and
    values were within normal limits. At autopsy, organ weights were
    measured and a microscopic examination made of the principal organs
    and tissues. No compound-related effects were observed (Olson &
    Voelker, 1970).

    Pig

         Groups of 8 Pitman-Moore miniature pigs were weaned at 3 days of
    age, and were fed formula diets containing 5.4% unmodified starch or
    4.9% hydroxypropyl distarch glycerol for 25 days. Growth was normal
    during the test period. At termination of the study, biochemical

    analyses of blood (haemoglobin) and serum (cholesterol, triglyceride,
    calcium, phosphorus, alkaline phosphatase, urea nitrogen, total
    protein, albumin and globulin) were similar for test and control
    animals. Relative organ weight as well as carcass composition (water,
    fat, protein, ash, Ca, PO4, Na, Mg) and liver composition (water,
    fat, protein and ash), were similar for test and control animals
    (Anderson et al., 1973a,b).

    Long-term studies

         Groups of 30 male and 30 female rats were fed for 104 weeks
    modified starch at 0, 5, 10 and 30% of their diet. No differences were
    noted between test and control animals with regard to appearance,
    behaviour, food consumption, haematology, serum chemistry and
    urinalysis. The mortality of females at the 30% level was slightly
    higher than in the control group. The growth rate was slightly reduced
    for both sexes at the 30% level when compared with controls. Caecal
    weight was increased at 30% in males and at 10% and 30% in females.
    Microscopic examination revealed no noticeable changes in the caecal
    wall. Renal calcification accompanied by focal hyperplasia of the
    pelvic epithelium was more marked in males fed 30% of the test
    compound than in control males. No distinct compound-related gross or
    microscopic changes were encountered in any of the organs examined (de
    Groot et al., 1974; Til et al., 1971a).

         Roe (1979) has conducted an extensive review of this type of
    mineral deposition in the renal pelvis of rats and concluded that
    pelvic nephrocalcinosis, corticomedullary nephrocalcinosis, acute
    tubular nephropathy, and calculus formation are manifestations of
    mineral imbalances and are of relatively common occurrence in
    untreated laboratory rats (particularly older animals).

    OBSERVATIONS IN MAN

         Twelve volunteers consumed 60 g modified starch on each of 4
    successive days. No adverse effects were noticed and no abnormalities
    occurred in regard to frequency or amount of faeces or faecal water
    and lactic acid content (Pieters et al., 1971).

    Comments

         The feeding studies with rats, dogs and pigs show that the
    modified starch is well utilized. The studies with radio-labelled
    hydroxypropyl starch provide some information in the rat on the
    metabolic fate of the hydroxypropyl moiety. Several short-term studies
    in the rat and dog show no adverse effects even at high dietary
    levels. The available evidence for the group of modified starches
    considered indicates that caecal enlargement without associated
    histopathological changes is without toxicological significance. The
    long-term and reproduction studies in the rat indicate no significant

    adverse effects and may be used for evaluation. The increased
    incidence of renal calcification accompanied by focal hyperplasia in
    test male rats in the long-term study is considered to be associated
    with imbalances in the Ca/P/ and Mg in the diet.

    EVALUATION

    Estimate of acceptable daily intake for man

    Not specified.*

              

    *    The statement "ADI not specified" means that, on the basis of the
         available data (toxicological, biochemical, and other), the total
         daily intake of the substance, arising from its use or uses at
         the levels necessary to achieve the desired effect and from its
         acceptable background in food, does not, in the opinion of the
         Committee, represent a hazard to health. For this reason, and for
         the reasons stated in individual evaluations, the establishment
         of an acceptable daily intake (ADI) in mg/kg bw is not deemed
         necessary.

    REFERENCES

    Anderson, T. A. et al. (1973a) Unpublished data submitted to Corn
         Refiners Ass., Inc.

    Anderson, T. A., Filer, L. J., Fomon, S. J., Andersen, D. W., Jensen,
         R. L. & Rogers, R. R. (1973b) Effect of waxy corn starch
         modification on growth, serum biochemical values and body
         composition of Pitman-Moore miniature pigs, Fd. Cosmet.
         Toxicol., 11, 747-754

    de Groot, A. P. & Spanjer, M. Th. (1970) Observations in rats fed on
         diets containing five different chemically modified starches.
         Report No. R 3096. Centraal Instituut voor Voedingsonderzoek,
         Zeist, Holland

    de Groot, A. P., Til, H. P., Feron, V. J., Van der Meullen, H. C. D.
         & Willems, M. I. (1974) Two-year feeding and multigeneration
         studies in rats on five chemically modified starches, Fd.
         Cosmet. Toxicol., 12, 651-664

    Knapp, W. A., Jr (1967a) Three-month dietary administration - rats,
         AR-197. Report of Hazleton Laboratories, Inc., Falls Church, Va.,
         to International Latex and Chemical Corporation, Hawthorne, N.J.
         Submitted to Federation of American Societies for Experimental
         Biology, Bethesda, Md., by A. E. Staley Manufacturing Company,
         Decatur, Ill.

    Knapp, W. A., Jr (1967b) 13-week dietary administration - dogs:
         AR-197 (test compound); tapioca fluor (reference compound).
         Report of Hazleton Laboratories, Inc., Falls Church, Va., to
         International Latex and Chemical Corporation, Hawthorne, N.J.
         Submitted to Federation of American Societies for Experimental
         Biology, Bethesda, Md., by A. E. Staley Manufacturing Company,
         Decatur, Ill.

    de Knecht-van Eekelen, A., Willems, M. & de Groot, A. P. (1970)
         Subchronic (90-day) toxicity study with a hydroxypropyl distarch
         glycerol in rats. Report No. R 3093. Centraal Instituut voor
         Voedingsonderzoek, Zeist, Holland

    Leegwater, D. C. (1971) Report No. 3431 by Centraal Instituut voor
         Voedingsonderzoek, Zeist, Holland. Submitted to WHO

    Olson, W. A. & Voelker, R. W. (1970) 13-week dietary administration -
         dogs: modified starch (F4-231); unmodified starch (control).
         Report of Hazleton Laboratories, Inc., Falls Church, Va.
         Submitted to A. E. Staley Manufacturing Company, Decatur, Ill.

    Olson, W. A. & Ferrell, J. F. (1970) Modified 13-week subacute one-
         generation reproduction study-rats. Unmodified starch. Modified
         starch (F4-231). Report of Hazleton Laboratories, Inc., Falls
         Church, Va. Submitted to A. E. Staley Manufacturing Company,
         Decatur, Ill.

    Pieters, J. J. L., van Staveren, W. A. & Brinkhuis, B. G. A. M. (1971)
         Report No. 3433 of the Centraal Instituut voor Voedingsonderzoek,
         Zeist, Holland. Submitted to WHO

    Roe, F. J. C. (1979) Mineral deposition in the renal pelvis of rats:
         A brief review, unpublished report. Submitted to WHO

    Til, H. P. et al. (1971a) Chronic (2-year) feeding study in rats with
         two chemically modified starches (starch acetate and
         hydroxypropyl distarch glycerol). Report No. R 3363. Centraal
         Instituut voor Voedingsonderzoek, Zeist, Holland

    Til, H. P., Spanjers, J. Th. & de Groot, A. P. (1971b) Unpublished
         report No. 3403 by Centraal Instituut voor Voedingsonderzoek,
         Zeist, Holland. Submitted to WHO
    


    See Also:
       Toxicological Abbreviations
       Hydroxypropyl distarch glycerol  (FAO Nutrition Meetings Report Series 46a)
       Hydroxypropyl distarch glycerol (WHO Food Additives Series 1)
       Hydroxypropyl distarch glycerol (WHO Food Additives Series 5)
       HYDROXYPROPYL DISTARCH GLYCEROL (JECFA Evaluation)