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    TRIAZOPHOS

    First draft prepared by Mr M. Watson,
    Ministry of Agriculture, Fisheries and Food,
    Harpenden, Hertfordshire, United Kingdom

    EXPLANATION

         Triazophos was evaluated for acceptable daily intake by the 1982
    JMPR (Annex I, 38) at which time a temporary ADI of 0-0.0002 mg/kg bw
    was established.  A toxicological monograph was published after that
    meeting (Annex I, 39).  Additional studies required to complete the
    toxicological data base were a carcinogenicity study, a teratology
    study in at least one mammalian species, metabolism studies in
    additional mammalian species (to explain species differences in acute
    toxicity studies) and mutagenicity studies.  Further studies of
    biochemical aspects, a range of mutagenicity studies and teratology
    studies in rats and rabbits were submitted for evaluation at the 1986
    Joint Meeting (Annex I, 47).  After consideration of these results the
    temporary ADI of 0-0.0002 mg/kg bw was extended but the final
    determination of an ADI was considered impractical since no
    carcinogenicity study was available.

         The results of carcinogenicity studies in rats and mice and
    additional studies that were submitted to the present meeting are
    summarized in this monograph addendum.

    EVALUATION FOR ACCEPTABLE INTAKE

    BIOLOGICAL DATA

    Biochemical aspects

    Effects on enzymes and other biochemical parameters

         Groups of 10 male and 10 female Wistar-derived rats were fed
    dietary levels of triazophos at 0, 3, 10 or 200 ppm for 48 weeks.  The
    treatment period was followed by a withdrawal period of 7 weeks during
    which all rats received untreated diets.  Cholinesterase activity in
    plasma and red blood cells was determined in all rats at weeks 33 and
    43 and during the withdrawal period at 3, 7, 14, 28 and 49 days.  At
    the end of the withdrawal period all rats were killed and
    cholinesterase activity in the brain was determined in 5 males and 5
    females from each group.  Cholinesterase activities in plasma and
    erythrocytes were distinctly inhibited at 10 ppm and considerably
    inhibited at 200 ppm.  Three days after discontinuing treatment with
    triazophos, the plasma cholinesterase activity in males had returned
    to normal.  In females (which were more severely affected during the
    treatment period), this recovery was observed within 7 days.  Recovery
    of cholinesterase activity in red blood cells took much longer. 
    Values were generally similar to controls after 2 to 4 weeks recovery
    at 10 ppm and after 4 to 7 weeks recovery at 200 ppm.  There were no
    inter-group differences in brain cholines-terase activity at
    termination (Til & Leegwater, 1974).

         The acute oral toxicity of triazophos (dissolved in arachis oil)
    was determined in groups of female rats, and the effects of different
    antidote therapies were investigated.  Antidotes were injected
    intraperitoneally, 1.5 minutes after dosing with triazophos. 
    Different antidotes (atropine sulphate, atropine methylnitrate,
    pralidoxime mesylate and obidoxime) were tested alone and in
    combination, but no attempt was made to investigate optimally
    effective doses of antidote.  From the results of this study it may be
    concluded that atropines raise the LD50 of triazophos by a factor of
    about 1.5.  The addition of pralidoxime to atropines does not increase
    the LD50 any further, but a combination of atropines and obidoxime
    raises the LD50 by a factor of about 2.  There was some indication
    that the toxic effects of triazophos are longer lasting than the
    antidote effects of atropines and oximes and that repeated dosing with
    antidotes may be beneficial, although this hypothesis was not tested
    directly in the experiment.  The study design of this experiment was
    inadequate to investigate the true effects of oxime antidotes on
    anti-cholinesterase poisoning induced by triazophos (Cohen, 1971).

    Toxicological studies

    Short-term studies

    Rats

         In a 4-week study, triazophos (purity 97.6%) was administered via
    inhalation exposure (nose only) to 4 groups of 10 male and 10 female
    Wistar rats at concentrations of 0 (Control), 1, 5 and 25 mg/m3 for
    6 hours per day, 5 days per week.  An additional 5 males and 5 females
    per group were used for a 4-week recovery period after completion of
    treatment.  There were no treatment-related clinical signs and the
    only death was associated with collection of a blood sample.  Body
    weight gain and food intake remained undisturbed by treatment and
    haematological and biochemical data apart from cholinesterase
    investigations revealed no changes of toxicological significance. 
    Depression of cholinesterase activity in plasma and erythrocytes was
    more marked in females than in males; males were affected only at the
    high dose, whereas females were affected at 5 and 25 mg/m3.  A
    treatment-related depression of brain cholinesterase activity was,
    however, seen only in males at the high dose.  Pathological
    investigations, including histopathology, revealed no indication of
    any reaction to treatment.  The depression of cholinesterase activity
    was completely reversible at the end of the 4-week recovery period. 
    The NOAEL was 5 mg/m3, based on the depression of
    acetylcholinesterase activity in brain at 25 mg/m3 (Bernstein
     et al., 1987).

         In a 4-week study, triazophos (purity 97.6%) was administered via
    dermal exposure (under an occlusive dressing) to 4 groups of 10 male
    and 10 female Wistar rats at dose levels of 0 (Control), 0.5, 5 and 50
    mg/kg bw for 6 hours per day, 5 days per week.  An additional 5 males
    and 5 females per group were used for a 4-week recovery period after
    completion of treatment.  There were no clinical signs of reaction to
    treatment, no treatment-related deaths and food intake and weight
    change remained unaffected.  Haematological investigations revealed no
    indication of any reaction to treatment.  A slightly increased glucose
    and urea level was noted for males receiving 5 mg/kg and both sexes
    receiving 50 mg/kg as well as decreased triglycerides and increased
    serum alanine aminotransferase activity for males receiving the high
    dose.  Cholinesterase activity was inhibited in plasma in females
    receiving 0.5 mg/kg bw/day and in both sexes at 5 and 50 mg/kg bw/day. 
    Erythrocyte cholinesterase was inhibited at 5 and 50 mg/kg bw/day in
    both sexes and brain acetylcholinesterase activity was inhibited only
    at the high dose level.  At pathological investigations, including
    histopathology, the only sign of reaction to treatment was a slight
    increase in adrenal weight in females treated with 5 or 50 mg/kg
    bw/day.  All of the changes seen during the treatment period were
    reversible during the withdrawal period.  The no observable adverse

    effect level was 0.5 mg/kg bw/day, based on the clinical chemistry and
    organ weight changes seen at 5 and 50 mg/kg bw/day (Thevenez  et al.,
    1987).

         In a 13-week dietary toxicity study, triazophos (purity 92.6%)
    was administered to groups of 10 male and 10 female Wistar rats at
    dietary concentrations of 0 (Control), 1, 20 and 400 ppm.  In order to
    investigate recovery from any reaction to treatment in a 4-week
    withdrawal period, an additional 10 animals per sex were added to the
    control, mid and high dietary level groups.  There were no clinical
    signs of reaction to treatment and no rats died prior to the scheduled
    necropsy.  A slight increase in food intake was noted for females
    receiving 400 ppm, although body weight gain remained unaffected by
    treatment.  Hematological investigations revealed evidence of a very
    slight degree of anaemia in rats treated with 400 ppm.  Investigation
    of cholinesterase activity revealed a dose-related depression in
    plasma and erythrocyte activity at 20 and 400 ppm and a depression in
    activity in brain at 400 ppm.  Pathological investigations at
    termination revealed no evidence of any reaction to treatment with
    triazophos.  All the changes seen during the treatment period were
    reversible during the withdrawal period.  The
    no-observable-adverse-effect-level was 20 ppm, equal to 1.5 mg/kg
    bw/day, based on the inhibition of brain cholinesterase activity at
    400 ppm (Tennekes  et al., 1986).

    Dogs

         In a 13-week study, groups of beagle dogs received triazophos
    (purity 92.6%) by dietary administration at levels of 0 (control),
    0.3, 9.0 and 270/180 ppm.  The control, mid- and high-dose groups
    consisted of 6 males and 6 females, while the low-dose group was
    composed of 4 dogs of each sex.  After completion of the treatment
    period, recovery from treatment-related effects was investigated in 2
    males and 2 females from the control and mid-dose group.  Severe
    cholinergic clinical signs of reaction to treatment were seen at the
    high dose.  Two males and one female from this group were killed in a
    moribund condition on days 43, 81 and 55, respectively.  The dietary
    level for the high-dose group was reduced from 270 to 180 ppm from day
    33 onwards.  Body weight loss and decreased food intake were seen at
    the high dose, but not at lower doses.  Haematological and biochemical
    changes were seen in dogs receiving 270/180 ppm but these were
    probably secondary to the poor general health of these animals.
    Inhibition of cholinesterase activity in plasma was seen at all dose
    levels, erythrocyte acetylcholinesterase was inhibited in dogs
    receiving 9.0 or 270/180 ppm and brain acetylcholinesterase was
    inhibited only in dogs receiving the high dose.  Pathological
    investigations revealed treatment-related changes only at the high
    dose.  These consisted of muscular hypertrophy in the duodenal wall,
    degenerative/inflammatory lesions in the zygomatic gland and
    mineralization of the fibrous-muscular tissue of the left ventricular

    papillary muscle.  Changes in cholinesterase activity in mid-dose dogs
    were reversible during the recovery period.  The NOAEL was 9.0 ppm,
    equal to 0.29 mg/kg bw/day, based on the severe effects, including
    mortality, seen at the high dose (Bloch  et al., 1986).

         In a 52-week study, groups of beagle dogs received triazophos by
    dietary administration at levels of 0, 0.2, 0.4, 4 or 80 ppm.  The
    control and high-dose groups consisted of 6 males and 6 females while
    the other groups were composed of 4 dogs of each sex.  Treatment with
    triazophos at 80 ppm was associated with the sacrifice of one female
    on day 73 and the termination of treatment in a second female from day
    106.  Both of these dogs showed signs typical of cholinesterase
    inhibition (persistent diarrhoea, evidence of dehydration, weight
    loss).  The withdrawal of treatment in the second female was followed
    by a progressive recovery, although cholinesterase activity remained
    below control and pre-test values.  Apart from the two females from
    the high-dose group mentioned above, diarrhoea was seen in most
    high-dose dogs and a few dogs receiving 4 ppm while evidence of
    dehydration and weight loss was seen in an additional male receiving
    4 ppm.  Investigations of auditory perception, ophthalmoscopy,
    haematological investigations and clinical chemistry tests including
    urinalysis but apart from cholinesterase investigations revealed no
    changes considered to be of any toxicological significance.  The
    assessment of plasma and erythrocyte cholinesterase activity indicated
    a dose-dependent inhibition for both sexes receiving 4 or 80 ppm. 
    Minor variations in cholinesterase activity at lower doses were
    considered not to be of toxicological significance and brain
    cholinesterase activity remained unaffected by treatment in all
    groups.  Pathological investigations at termination of the study,
    including autopsy findings, organ weight analysis and
    histopathological examination of tissues revealed no evidence of any
    reaction to treatment with triazophos. The NOAEL was 4 ppm (equal to
    0.12 mg/kg bw/day) based on inhibition of cholinesterase activity and
    signs of reaction to treatment, including mortality, at 80 ppm (Allen
     et al., 1989).

    Monkeys

         Groups of one male and one female adult rhesus monkeys were given
    daily oral doses of triazophos (suspended in starch slurry) of 0.025
    or 0.05 mg/kg bw by stomach intubation for 22 consecutive days. 
    Behaviour, general condition, body weight and food intake of the
    animals was monitored.  Blood samples were collected for determination
    of serum and erythrocyte cholinesterase activity pre-test, after each
    dose and twice weekly after the last dose for 6 weeks.  Behaviour and
    general condition of the animals was not affected by treatment and
    food consumption was affected only in the male monkey receiving the
    high dose.  This animal ate 40-45% less than expected during the
    treatment period, but intake returned quickly to normal after
    cessation of dosing.  Body weight remained unaffected by treatment in

    all animals.  Serum cholinesterase activity in animals receiving
    0.05 mg/kg bw/day fell rapidly to about 30-35% of the initial value,
    with the maximum decrease being seen after 5-8 days dosing.  After the
    last dose, activities increased continuously, approaching the initial
    value after 18-22 days withdrawal.  At the lower dose, serum
    cholinesterase activity fell to about 40% of initial activity in the
    female and 70% in the male.  Lowest values were seen after about 18
    days dosing and values increased after the end of the dosing period;
    reaching initial values after 8 days withdrawal in the male and 15
    days in the female.  Erythrocyte cholinesterase activity remained
    unaffected by treatment at either 0.025 or 0.05 mg/kg bw/day (Scholz
    & Baeder, 1971).

    Long-term/carcinogenicity studies

    Mice

         In a combined long-term toxicity and carcinogenicity study in
    mice, triazophos was administered via the diet to four groups of 60
    male and 60 female NMRI mice at dietary levels of 0, 6, 30 or 150 ppm. 
    From each group 10 males and 10 females were designated for interim
    sacrifice after 52 weeks and the remaining 50 males and 50 females
    comprised the carcinogenicity study and were scheduled for sacrifice
    after 104 weeks.

         There were no clinical signs of reaction to treatment and
    although mortality at the high dose was slightly higher than in
    controls, this difference did not attain statistical significance. 
    Survival at the terminal sacrifice was in the range of 46 - 52% in the
    control, 6 and 30 ppm groups and 38% in the high dietary level group. 
    Food intake and weight gain remained largely unaffected by treatment
    and hematological parameters also remained unaffected by treatment
    with triazophos.  Clinical chemistry investigations revealed
    treatment-related changes in cholesterol levels and cholinesterase
    activity.  At the 12-month interim sacrifice serum cholesterol levels
    were increased, compared to controls in high dose male mice; a
    similar, though not statistically significant, change was seen at the
    terminal sacrifice in males treated with 30 or 150 ppm.  At the
    interim sacrifice, serum cholinesterase activity was inhibited at 30
    and 150 ppm, and erythrocyte cholines-terase was inhibited in males at
    150 ppm and in females at 30 and 150 ppm.  More severe effects were
    seen at the terminal sacrifice: serum and erythrocyte cholinesterase
    were inhibited, compared to controls, in all groups, apart from
    erythrocyte activity in males treated with the low dose.  Brain
    cholinesterase activity was not affected by treatment after 52 weeks
    but at the terminal sacrifice a marked inhibition was seen in females
    treated with 150 ppm, though not in any other group.  Pathological
    investigations at the interim and terminal sacrifices revealed no
    indication of any reaction to treatment with triazophos.  In
    particular, the type and incidence of neoplastic lesions were those

    commonly seen in aged mice.  The NOAEL was thus 30 ppm, equal to
    4.5 mg/kg bw/day, based on the slightly increased mortality seen in
    animals treated with 150 ppm and inhibition of brain cholinesterase in
    females at 150 ppm (Donaubauer  et al., 1989).

    Rats

         In a combined long-term toxicity and carcinogenicity study in
    rats, triazophos was administered via the diet to four groups of 80
    male and 80 female Wistar-derived rats at dietary levels of 0, 3, 27,
    or 243 ppm.  From each group, 10 males and 10 females were designated
    for interim sacrifice after 52 weeks, 20 males and 20 females for
    sacrifice after 104 weeks, while the remaining 50 males and 50 females
    in each group comprised the carcinogenicity study and were scheduled
    for sacrifice after 118 weeks of treatment.  There were no clinical
    signs of reaction to treatment and mortality remained unaffected by
    administration of triazophos.  Survival at the terminal sacrifice was
    in the range of 46-51% in the different groups.  Female rats in the
    high dose group ate about 10% more food than control rats and this was
    reflected in an increased body weight gain in this group; otherwise
    appetite and growth remained unaffected by treatment.  In general,
    treated females tended to drink more water than did the controls, this
    effect was most evident at the high dietary level but males remained
    unaffected.  Hearing tests and ophthalmoscopy revealed no
    treatment-related changes.  Haematology and urinalysis investigations
    revealed no indication of any reaction to treatment and
    treatment-related changes in clinical biochemistry tests were confined
    to the inhibition of cholinesterase activity.  There was a
    dose-related inhibition of cholinesterase activity in plasma and
    erythrocytes at 243 and 27 ppm during the course of treatment.  In
    females treated with 243 ppm a slight (approximately 21-28%)
    inhibition of brain cholinesterase activity was noted at both the
    interim (52 week) and chronic toxicity (104 weeks) sacrifices. 
    Post-mortem examinations revealed no indication of any effect on organ
    weights but there was a higher incidence of nodular changes in the
    pancreas and small intestine in males treated with 243 ppm than in
    controls.  This correlated with an increased incidence of focal or
    multifocal hyperplasia of the exocrine pancreas in treated groups. 
    This change was seen in 9 males at 243 ppm, 5 males at 27 ppm and 1
    male at 3 ppm, but not in any controls.  This lesion was not found in
    any female rats from any group.  The incidence and distribution of
    tumours remained unaffected by treatment; all the neoplastic lesions
    encountered were of types frequently seen in the strain of rat used in
    this study.  The NOAEL was thus 3 ppm, equal to 0.17 mg/kg bw/day
    based on the increased incidence of hyperplasia of the exocrine
    pancreas observed at 243 and 27 ppm in addition to inhibition of brain
    cholinesterase in females at 243 ppm (Tennekes  et al., 1990).

    Reproduction studies

         In a two-generation (one litter per generation) reproduction
    study, groups of 25 male and 25 female Wistar rats received dietary
    administration of triazophos at 0, 3, 27 or 243 ppm.  Dietary levels
    were based on the results of a preliminary study (Suter & Terrier,
    1989).  Treatment was continued for a 70 day premating period, and
    throughout pairing, gestation and lactation for breeding of the F1
    litters.  Following weaning on day 21 post-partum, groups of 25 male
    and 25 female F1 animals were selected.  Diets were fed to these
    animals for 125 days prior to pairing, and throughout pairing,
    gestation and lactation for breeding of the F2 litters.  Clinical
    signs of reaction to treatment were seen at 243 ppm.  Aggressive
    behaviour was noted in the parental generation during the premating
    period; exophthalmia, ataxia, tremors and dyspnoea were noted during
    the lactation period in the females of the parental and F1
    generations and in F1 pups for up to 3 weeks at the start of the
    premating period.  Three F1 generation females receiving 243 ppm died
    during the lactation period.  Treatment-related reductions in food
    intake and weight gain were noted at 243 ppm, and a higher breeding
    loss was also seen at the high dietary level than in the control
    group.  Reproductive index and general fertility parameters remained
    unaffected by treatment at all dietary levels and there was no
    evidence of any obvious teratogenic effect.  Organ weights and
    pathological examination gave no indication of any reaction to
    treatment and behaviour, food intake and weight gain and pup viability
    remained unaffected by treatment at 3 or 27 ppm.  The NOAEL was thus
    27 ppm, equivalent to 2-3 mg/kg bw/day (Suter  et al., 1989).

    Special studies on delayed neurotoxicity

         An acute delayed neurotoxicity study was performed in hens. 
    Preliminary investigations of the acute oral toxicity revealed that
    the LD50 in white Leghorn hens was 7.5 (4.8-10.1) mg/kg bw, and that
    protection with atropine sulfate and 2-PAM methiodide raised the LD50
    to approximately 50 mg/kg.  In the main study, triazophos was
    administered twice, orally, at the protected LD50 dose of 50 mg/kg bw
    to a group of 20 white Leghorn hens, with an interval of 21 days
    between doses.  A negative control group and a positive control group
    (TOCP 500 mg/kg bw), each composed of 6 animals, were also included in
    the study.  Dosing with triazophos resulted in the death of 15 animals
    and the sacrifice of one further animal on day 38, despite symptomatic
    treatment with antidotes.  Deaths occurred within 24 hours of dosing,
    9/20 after the first dose and a further 6 after the second.  Of the
    four hens which survived to the end of the study, two showed slight to
    severe signs of ataxia.  In one animal signs were seen 7 days after
    the second dose and in the other 12 days after the second dose.  These
    clinical signs were accompanied by histological lesions in the spinal
    cord (cystic dilatation in myelin sheaths and plaque-shaped

    decomposition bodies).  Similar histological lesions were also present
    in the hen which showed no ataxia but was killed on day 38.  Animals
    in the positive control group showed the typical neurotoxic effects
    produced by TOCP (Ebert & Mayer, 1988).

         In a further acute delayed neurotoxicity test, triazophos was
    administered twice at a dose level of 12.5 mg/kg bw to a group of 15
    white Leghorn hens protected by an antidote treatment with an interval
    of 21 days between doses.  A negative control group and a positive
    control group (TOCP 500 mg/kg bw), each composed of 6 animals, were
    also included in the study.  All hens treated with triazophos survived
    and were killed at the end of the study.  Cholinergic symptoms such as
    lacrimation and trembling were seen shortly after dosing with
    triazophos.  Assessment of ataxia and histological examination of
    brain, spinal cord and peripheral nerves indicated no neurotoxic
    effects of triazophos.  In contrast, animals in the positive control
    group, treated with TOCP, showed the typical neurotoxic effects such
    as severe delayed ataxia or paralysis and histological lesions such as
    axonal swellings in the spinal cord and decomposition of the myelin
    sheaths together with reactive proliferation of the Schwann's cells in
    peripheral nerves (Ebert, 1989).

         Neurotoxicity has also been investigated via short term, dietary
    administration of triazophos. In a 20-day preliminary study,
    triazophos was administered to groups of five female white Leghorn
    hens by dietary admixture, at levels of 0 (control), 50, 100, 150 or
    200 ppm.  At termination, three animals from each group were assigned
    for pathology (preservation of tissues only) and two for determination
    of neuropathy target esterase (NTE) activity.  All animals survived
    until termination, there were no clinical signs of reaction to
    treatment, and food intake, weight change, forced motor activity and
    NTE activity remained undisturbed by treatment.

         In the main study, triazophos was administered to groups of 10
    female, white Leghorn hens by dietary admixture, at levels of 0
    (control), 50, 110 or 250 ppm.  There were no treatment-related
    clinical signs among animals receiving 50 or 110 ppm.  At 250 ppm,
    restless and/or excited behaviour was seen in all animals and sedation
    in 2/10.  One of these latter two hens died on day 76.  At 50 and 110
    ppm food intake was initially reduced compared to controls but was
    unaffected overall.  At 250 ppm food intake was consistently reduced. 
    Treated animals displayed a dose-related weight loss during the study. 
    Examination of forced motor activity revealed no treatment-related
    changes at 50 or 110 ppm, while at 250 ppm, two of the ten animals
    displayed symptoms characteristic of delayed neurotoxicity.  One of
    these died on day 76.  Plasma cholinesterase activity was markedly
    reduced at all dose levels.  NTE activity was not investigated. 
    Patho-logical examination revealed no treatment-related macroscopic
    changes.  Histo-logical examination revealed morphological evidence of
    short-term neuro-toxicity in the spinal cord and peripheral nerves in
    animals treated with 250 ppm.  Similar lesions were seen in control

    and treated hens, but the severity of the lesions and the number of
    animals affected was greater at the high dietary level than in the
    other groups (Ullmann  et al., 1991).

    Special studies on skin sensitization and irritation

         The dermal sensitization potential of triazophos has been tested
    in guinea pigs according to the Buehler patch test method.  The
    induction phase was carried out by repeated dermal application of a
    10% solution of triazophos in sesame oil.  At challenge (using a 5%
    solution) no dermal reaction was seen and it was concluded that
    triazophos displayed no sensitizing potential in this test (Hollander
    & Weigand, 1976).

    COMMENTS

         In 1986 the temporary ADI of 0-0.0002 mg/kg bw was extended
    pending the submission of a carcinogenicity study.

         In a 52-week study in dogs, employing dietary concentrations of
    0, 0.2, 0.4, 4, or 80 ppm triazophos, the only signs of reaction to
    treatment were associated with inhibition of plasma and erythrocyte
    cholinesterase activity.  The NOAEL was 4 ppm (equal to 0.12 mg/kg
    bw/day), based on inhibition of plasma and erythrocyte cholinesterase
    activity and mortality at 80 ppm.

         Long-term/carcinogenicity studies in rats and mice at dietary
    concentrations of 0, 3, 27, or 243 ppm and 0, 6, 30 or 150 ppm
    demonstrated that triazophos has no carcinogenic potential in either
    species.  In rats an increased incidence of hyperplasia of the
    exocrine pancreas was observed at 27 and 243 ppm and inhibition of
    brain acetylcholinesterase in females at 243 ppm.  The NOAEL was 3
    ppm, equal to 0.17 mg/kg bw/day.  In mice, slightly increased
    mortality at 150 ppm and inhibition of brain acetylcholinesterase in
    females at 150 ppm were observed.  The NOAEL was 30 ppm, equal to
    4.5 mg/kg bw/day.

         In a two-generation reproduction study in rats at dietary
    concentrations of 3, 27 or 243 ppm triazophos, marked clinical signs,
    including mortality, decreased survival and reduced weight gain in
    pups were seen at 243 ppm.  The NOAEL was 27 ppm, equal to
    2-3 mg/kg bw/day.

         There were two additional delayed neurotoxicity tests in hens. 
    In the first, atropine and oxime antidotes were used with a dose of
    triazophos about 7 times the unprotected LD50 of 7.5 mg/kg bw.  In the
    group treated with triazophos, high mortality was seen and only 4 hens
    survived to termination.  Among these 4 hens, signs of ataxia were
    seen in 2 hens and histological lesions were seen in the nervous
    tissue of 3 animals.  In the second study, at lower doses (about twice
    the unprotected LD50), triazophos displayed no neurotoxic effects.

         A 90-day neurotoxicity study in hens at dietary concentrations of
    0, 50, 110 or 250 ppm, was preceded by a 20-day preliminary study.  In
    the preliminary study, at dietary levels up to 200 ppm, there were no
    neurotoxic symptoms and neuropathy target esterase (NTE) activity
    remained undisturbed by treatment.  Histopathology was not performed. 
    In the main study, cholinergic signs were seen at the high dose of 250
    ppm and one hen (of 10) died on day 76.  This hen, and one other,
    displayed clinical signs of neurotoxicity.  Plasma cholinesterase
    activity was markedly reduced at all doses, but NTE activity was not
    investigated.  Morphological lesions characteristic of neurotoxicity
    were reported in the spinal cord and peripheral nerves.  The study
    report stated that lesions were more severe and seen more frequently

    in hens treated with triazophos than in controls.  The Meeting
    concluded that it was difficult to interpret these histopathology
    findings since, most unusually, lesions were frequently seen in the
    spinal cord, but very infrequently in the peripheral nerves, of
    control animals.

         The Meeting reviewed the results of male and female human
    volunteer studies previously summarized in the 1982 monograph.  A
    series of experiments had been conducted, including a three-week trial
    with 25 volunteers.  The present Meeting agreed with the conclusion of
    the 1982 JMPR regarding that trial: a dose of 0.0125 mg/kg bw/day of
    triazophos was a minimal effect level with regard to plasma
    cholinesterase activity but was without effect on erythrocyte
    acetylcholin-esterase.

         In view of the uncertainty regarding the potential for triazophos
    to cause delayed neurotoxicity, the Meeting extended the temporary
    ADI, basing it on the NOAEL in dogs, utilizing a 500-fold safety
    factor.

    TOXICOLOGICAL EVALUATION

    Level causing no toxicological effect

         Mouse:    30 ppm in the diet, equal to 4.5 mg/kg bw/day
         Rat:      3 ppm in the diet, equal to 0.17 mg/kg bw/day
                   (long term study)
                   27 ppm in the diet, equal to 2-3 mg/kg bw/day
                   (multigeneration study)
         Dog:      4 ppm in the diet, equal to 0.12 mg/kg bw/day
         Human:    0.0125 mg/kg bw/day

    Estimate of temporary acceptable daily intake for humans

         0-0.0002 mg/kg bw

    Studies without which the determination of a full ADI is
    impractical

         To be submitted to WHO by 1992:

    1.   Clarification of the potential for triazophos to cause delayed
         neurotoxicity through an acute delayed neurotoxicity study in
         hens combined with measurements of acetylcholinesterase and
         neuropathy target esterase inhibition in nervous tissue.

    2.   Submission of ongoing studies, including those on antidotes to
         acute poisoning.

    REFERENCES

    Allen, T.R., Frei, T., Luetkemeier, H., Vogel, W., Terrier, Ch.,
    Vogel, O. & Armstrong, J. (1989)  Triazophos technical grade: Chronic
    oral toxicity, 52-week feeding study in beagle dogs.  Unpublished
    report from RCC, submitted by Hoechst AG., Frankfurt-am-Main, Germany.

    Bernstein, D.M., Luetkemeier, H., Vogel, W., Schlotke, B., & Terrier,
    Ch. (1987)  Triazophos technical grade: Subacute 28-day repeated dose
    inhalation toxicity study in rats. Unpublished report from RCC,
    submitted by Hoechst AG., Frankfurt-am-Main, Germany.

    Bloch, I., Frei, Th., Luetkemeier, H., Vogel, W. & Terrier, Ch. (1986) 
    Triazophos technical grade: subchronic oral toxicity 13-week feeding
    study in beagle dogs. Unpublished report from RCC, submitted by
    Hoechst AG., Frankfurt-am-Main, Germany.

    Cohen, E.M. (1971)  Therapeutic effects of various antidotes against
    intoxication with the compound Hoe 02960 in rats. Unpublished report
    from TNO submitted by Hoechst AG., Frankfurt-am-Main, Germany.

    Donauber, H.H. (1989)  Triazophos substance technical: Combined
    chronic toxicity and carcinogenicity study in mice. Unpublished report
    from Hoechst AG., Frankfurt-am-Main, Germany.

    Ebert, E. (1989)  Triazophos substance technical:  Testing for acute
    delayed neurotoxicity in white leghorn hens. Unpublished report from
    Hoechst AG., Frankfurt-am-Main, Germany.

    Ebert, E. and Mayer, D. (1988)  Triazophos substance technical: 
    Testing for acute delayed neurotoxicity in white leghorn hens. 
    Unpublished report from Hoechst AG., Frankfurt-am-Main, Germany.

    Hollander & Weigand (1976)  Testing for sensitizing properties of
    triazophos in the guinea pig.  Unpublished report from Hoechst AG.,
    Frankfurt-am-Main, Germany.

    Scholz, J. & Baeder, C. (1971)  Report on a 22-day feeding trial with
    Hoe 2960 and rhesus monkeys.  Unpublished report from Hoechst AG.,
    Frankfurt-am-Main, Germany.

    Suter, P., Vogel, W., Armstrong, J.M. & Terrier, Ch. (1989) 
    Triazophos technical grade:  The two-generation reproduction study in
    the rat.  Unpublished report from RCC, submitted by Hoechst AG.,
    Frankfurt-am-Main, Germany.

    Suter, P., Vogel, W. & Terrier, Ch. (1989)  Triazophos technical
    grade:  Preliminary study to the two-generation reproduction study in
    the rat.  Unpublished report from RCC, submitted by Hoechst AG.,
    Frankfurt-am-Main, Germany.

    Tennekes, H., Janiak, T., Probst, D., Leutkemeier, H., Vogel, O.,
    Westen, H., Biedermann, K. & Heusner, W. (1990)  Triazophos technical
    grade:  Chronic toxicity/oncogenicity feeding study in rats.
    Unpublished report from RCC, submitted by Hoechst AG.,
    Frankfurt-am-Main, Germany.

    Tennekes, H., Horst, K., Luetkmeier, H., Vogel, W., Schlotke, B. &
    Terrier, Ch. (1986)  Triazophos technical grade: subchronic oral
    toxicity 13 week feeding study in rats. Unpublished report from RCC,
    submitted by Hoechst AG., Frankfurt-am-Main, Germany.

    Thevenez, Th., Luetkemeier, H., Schlotke, B., Vogel, W. & Terrier, Ch.
    (1987)  Triazophos technical grade: subacute 28-day repeat dose dermal
    toxicity study in rats. Unpublished report from RCC, submitted by
    Hoechst AG., Frankfurt-am-Main, Germany.

    Til, H.P. and Leegwater, D.C. (1974)  Investigation of the recovery of
    cholinesterase inhibition induced by Hoe 2960 in rats.  Unpublished
    report from CIVO/TNO, submitted by Hoechst AG., Frankfurt-am-Main,
    Germany.

    Ullmann, L., Sacher, R., Porricello, T., Janiak, Th., Luetkemeier, H.,
    Biedermann, K., Vogel, O., Terrier, Ch.  (1991) 3-Month subchronic
    delayed neurotoxicity (feeding) study with triazophos - substance
    technical in the hen. RCC Report 123214 of 13 June, submitted by
    Hoechst AG., Frankfurt-am-Main, Germany.


    See Also:
       Toxicological Abbreviations
       Triazophos (JMPR Evaluations 2002 Part II Toxicological)
       Triazophos (Pesticide residues in food: 1982 evaluations)
       Triazophos (Pesticide residues in food: 1983 evaluations)
       Triazophos (Pesticide residues in food: 1986 evaluations Part II Toxicology)
       Triazophos (Pesticide residues in food: 1993 evaluations Part II Toxicology)